But isn't it enough that the gene for expressing such elaborate displays is 'selfish'? Consider that a gene that 'wants to be copied' (all genes), if it could signal it's presence and simultaneously cause that signal to trigger mating, would succeed in continuing in the species? In that case mating rituals are not there for any fitness proof, but simply a sort of parasite. A gene that's using an animal for it's own purposes.
How could we even distinguish this sort of parasitic gene, from any other kind of fitness selection? Is there a meaningful difference?
DNA is an inert chemical chain encoded with linear and non linear data which is acted upon and maintained by nano machines manufactured with the instructions it contains, which do you think came first?